In Elizabethan London, Matthew and Diana's romance faces a barrage of new threats. Diana's unleashed magic takes a dark and frightening turn, while Matthew struggles to re-inhabit the dangerous life he led over four centuries ago. They must overcome deep personal fears and jealousies, baring their darkest secrets to one another if they are to stay alive, stay together and find a way back to present day. While Matthew and Diana hide out in Elizabethan London, back in present day, Diana's beloved aunts, Sarah and Em, must take shelter with notorious witch hunter Ysabeau de Clermont at her ancestral home, Sept-Tours. Meanwhile, in Oxford, Marcus and Miriam take on Matthew's mantle to protect daemons Nathaniel and Sophie, whose pregnancy is advancing. And Gerbert, Knox, Satu and Domenico are determined to hunt down every clue they can to Diana's and Matthew's disappearance, and the secrets their allies are keeping from them.